I see this as similar to the Lucifer/Judas-like understanding that such rebellions and betrayals of God/The Increate are part of the Plan and necessary to it. This tiny Tzadkiel has been broken off from the larger being and, apprently, she was rebellious against the larger angel and has been banished to this realm for doing that. I interpret this disconnect between Typhon and the megatherians in light of the tiny Tzadkiel angel that Severian encounters at the Brook Madregot. Typhon assures Severian that these beings will soon be his abject slaves. But when Typhon wakes up and is discussing the current state of Urth with Severian, he is told that the world is ruled by gigantic sea beings (one of whom is Scylla, as mentioned by the Caloyer of Saltus). Mythologically, and in Long/Short Sun, Scylla is the daughter of Typhon. I'll risk over-dissecting the enigmatic nature of these beings to note that the text probably doesn't allow us to consider megatherians to be Typhon's pets exactly.
Each view being an alternate perspective of the same thing. I think Wolfe was highly influenced by Lovecraft and I think he might consider the Cthulhu Mythos to be another way of perceiving our spiritual universe, cognate rather than separate from Biblical, Greek, Hindu and Zoroastrian cosmologies. That's a refreshing take that I can appreciate.
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